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-25.1146 -1.3712 Td [(51.)-499.6 (Zhaxybayeva)-347.6 (O,)-347.6 (Gogarten)-347.6 (JP)-347.6 (\(2003\))-347.6 (An)-347.6 (improved)-347.6 (probability)-347.6 (mapping)-347.6 (approach)-347.6 (to)]TJ 1.8897 -1.3712 Td [(assess)-288.9 (genome)-288.9 (m)-0.1 (osaicism.)]TJ /T1_1 1 Tf 12.3702 0 Td [(BMC)-288.9 (Genomics)]TJ /T1_0 1 Tf 7.2441 0 Td (4:37.)Tj -21.5039 -1.3712 Td [(52.)-499.6 (Doolittle)-291.7 (WF)-291.7 (\(1999\))-291.7 (Lateral)-291.7 (genomics.)]TJ /T1_1 1 Tf 19.742 0 Td [(Trends)-291.7 (Cell)-291.7 (Biol)]TJ /T1_0 1 Tf 7.4884 0 Td (9:M5\226M8.)Tj -27.2304 -1.3712 Td [(53.)-499.6 (Ochman)-250 (H,)-250 (Lawrence)-250 (JG,)-250 (Groisman)-250 (EA)-250 (\(2000\))-250 (Lateral)-250 (gene)-250 (transfer)-250 (and)-250 (the)-250 (nature)-250 (of)]TJ 1.8897 -1.3712 Td [(bacterial)-278 (innovation.)]TJ /T1_1 1 Tf 9.9474 0 Td (Nature)Tj /T1_0 1 Tf 3.5011 0 Td [(405:299)-119.8 (\226304.)]TJ -15.3383 -1.3712 Td [(54.)-499.6 (Gogarten)-261.9 (JP,)-261.9 (Doolittle)-261.9 (WF,)-261.9 (Lawrence)-261.9 (JG)-261.9 (\(2002\))-261.9 (Prokaryotic)-261.9 (evolution)-261.9 (in)-261.9 (light)-261.9 (of)-261.9 (gene)]TJ 1.8897 -1.3712 Td (transfer.)Tj /T1_1 1 Tf 4.2516 0 Td [(Mol)-305.4 (Biol)-305.4 (Evol)]TJ /T1_0 1 Tf 6.4725 0 Td [(19:2226)-119.8 (\2262238.)]TJ -12.6139 -1.3712 Td [(55.)-499.6 (Shimodaira)-516.1 (H,)-516.1 (Hasegawa)-516.1 (M)-516.2 (\(1999\))-516.1 (Multiple)-516.1 (comparisons)-516.1 (of)-516.1 (log-likelihoods)-516.1 (with)]TJ 1.8897 -1.3712 Td [(applications)-293.6 (to)-293.6 (phylogenetic)-293.6 (inference.)]TJ /T1_1 1 Tf 18.5113 0 Td [(Mol)-293.6 (Biol)-293.6 (Evol)]TJ /T1_0 1 Tf 6.437 0 Td [(16:1114)-119.8 (\2261116.)]TJ -26.838 -1.3712 Td [(56.)-499.6 (Race)-396.8 (HL,)-396.8 (Herrmann)-396.8 (RG,)-396.8 (Martin)-396.8 (W)-396.8 (\(1999\))-396.8 (Why)-396.8 (have)-396.8 (organelles)-396.8 (retained)-396.8 (genomes?)]TJ /T1_1 1 Tf 1.8897 -1.3712 Td [(Trends)-305.4 (Genet)]TJ /T1_0 1 Tf 6.5572 0 Td [(15:364)-119.8 (\226370.)]TJ -8.4468 -1.3712 Td [(57.)-499.6 (Martin)-486.3 (W,)]TJ /T1_1 1 Tf 7.3075 0 Td [(et)-486.3 (al.)]TJ /T1_0 1 Tf 3.0297 0 Td [(\(2002\))-486.3 (Evolutionary)-486.3 (analysis)-486.3 (of)]TJ /T1_1 1 Tf 15.2829 0 Td (Arabidopsis)Tj /T1_0 1 Tf [-0.3 (,)-486.3 (cyanobacterial,)-486.3 (and)]TJ -23.7304 -1.3712 Td [(chloroplast)-176.6 (genomes)-176.6 (reveals)-176.6 (plastid)-176.7 (phylogeny)-176.6 (and)-176.6 (thousands)-176.6 (of)-176.6 (cyanobacterial)-176.6 (genes)]TJ 0 -1.3712 TD [(in)-305.4 (the)-305.4 (nucleus.)]TJ /T1_1 1 Tf 7.1396 0 Td [(Proc)-305.4 (Natl)-305.4 (Acad)-305.4 (Sci)-305.4 (USA)]TJ /T1_0 1 Tf 10.9716 0 Td [(99:12246)-119.8 (\22612251.)]TJ -20.0008 -1.3712 Td [(58.)-499.6 (Shi)-194.6 (T,)-194.6 (Bibby)-194.6 (TS,)-194.6 (Jiang)-194.6 (L,)-194.6 (Irwin)-194.6 (AJ,)-194.6 (Falkowski)-194.6 (PG)-194.6 (\(2005\))-194.6 (Protein)-194.6 (interactions)-194.6 (limit)-194.6 (the)-194.6 (rate)]TJ 1.8897 -1.3712 Td [(of)-288.8 (evolution)-288.9 (of)-288.8 (photosynthetic)-288.9 (genes)-288.8 (in)-288.9 (cyanobacteria.)]TJ /T1_1 1 Tf 25.8592 0 Td [(Mol)-288.8 (Biol)-288.8 (Evol)]TJ /T1_0 1 Tf 6.4228 0 Td [(22:2179)-119.8 (\2262189.)]TJ -34.1717 -1.3712 Td [(59.)-499.6 (Dandekar)-176.6 (T,)-176.6 (Snel)-176.6 (B,)-176.6 (Huynen)-176.7 (M,)-176.6 (Bork)-176.6 (P)-176.7 (\(1998\))-176.6 (Conservation)-176.6 (of)-176.6 (gene)-176.6 (order:)-176.6 (A)-176.6 (\036ngerprint)]TJ 1.8897 -1.3712 Td [(of)-291.7 (proteins)-291.7 (that)-291.7 (physically)-291.7 (interact.)]TJ /T1_1 1 Tf 16.5733 0 Td [(Trends)-291.7 (Biochem)-291.7 (Sci)]TJ /T1_0 1 Tf 9.2095 0 Td [(23:324)-119.8 (\226328.)]TJ -27.6724 -1.3712 Td [(60.)-499.6 (Blankenship)-203.2 (RE)-203.2 (\(2001\))-203.1 (Molecular)-203.1 (evidence)-203.2 (for)-203.2 (the)-203.2 (evolution)-203.2 (of)-203.1 (photosynthesis.)]TJ /T1_1 1 Tf 38.04 0 Td (Trends)Tj -36.1503 -1.3712 Td [(Plants)-305.4 (Sci)]TJ /T1_0 1 Tf 0.1198 Tc 4.612 0 Td [(6)119.8 (:)119.8 (4\2266)119.8 (.)]TJ 0 Tc -6.5017 -1.3712 Td [(61.)-499.6 (Fennel)-177.6 (K,)-177.6 (Follows)-177.6 (M,)-177.6 (Falkowski)-177.6 (PG)-177.6 (\(2005\))-177.6 (The)-177.6 (co-evolution)-177.6 (of)-177.6 (the)-177.6 (nitrogen,)-177.6 (carbon)-177.6 (and)]TJ 1.8897 -1.3712 Td [(oxygen)-289.8 (cycles)-289.8 (in)-289.8 (the)-289.8 (Proterozoic)-289.8 (ocean.)]TJ /T1_1 1 Tf 18.5747 0 Td [(Am)-289.8 (J)-289.8 (Sci)]TJ /T1_0 1 Tf 4.0915 0 Td [(305:526)-119.8 (\226545.)]TJ -24.5559 -1.3712 Td [(62.)-499.6 (Anbar)-200.3 (AD,)-200.3 (Knoll)-200.3 (AH)-200.3 (\(2002\))-200.3 (Proterozoic)-200.3 (ocean)-200.3 (chemistry)-200.3 (and)-200.3 (evolution:)-200.3 (A)-200.3 (bioinorganic)]TJ 1.8897 -1.3712 Td (bridge?)Tj /T1_1 1 Tf 3.8342 0 Td (Science)Tj /T1_0 1 Tf 3.6671 0 Td (297:1137\2261142.)Tj -9.391 -1.3712 Td [(63.)-499.6 (Raymond)-164.8 (J,)-164.8 (Siefert)-164.8 (JL,)-164.8 (Staples)-164.8 (CR,)-164.8 (Blankenship)-164.8 (RE)-164.8 (\(2004\))-164.8 (The)-164.8 (natural)-164.8 (history)-164.8 (of)-164.8 (nitrogen)]TJ 1.8897 -1.3712 Td (\036xation.)Tj /T1_1 1 Tf 4.1956 0 Td 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-1.3712 Td [(66.)-499.6 (Tatusov)-487.2 (RL,)]TJ /T1_1 1 Tf 7.8654 0 Td [(et)-487.2 (al.)]TJ /T1_0 1 Tf 3.0316 0 Td [(\(2001\))-487.2 (The)-487.3 (COG)-487.3 (database:)-487.2 (New)-487.3 (developments)-487.2 (in)-487.3 (phylogenetic)]TJ -9.0073 -1.3712 Td [(classi\036cation)-289.8 (of)-289.8 (proteins)-289.8 (from)-289.8 (complete)-289.8 (genomes.)]TJ /T1_1 1 Tf 23.577 0 Td [(Nucl)-289.8 (Acids)-289.8 (Res)]TJ /T1_0 1 Tf 6.9247 0 Td (29:22\22628.)Tj -32.3913 -1.3712 Td [(67.)-499.6 (Thompson)-438 (J,)-438 (Higgins)-438 (D,)-438 (Gibson)-438 (T)-438 (\(1994\))-438 (CLUSTAL)-438 (W:)-438 (Improving)-438 (the)-438 (sensitivity)-438 (of)]TJ 1.8897 -1.3712 Td [(progressive)-516.6 (multiple)-516.6 (sequence)-516.6 (alignment)-516.6 (through)-516.6 (sequence)-516.6 (weighting,)-516.6 (position-)]TJ T* [(speci\036c)-288.8 (gap)-288.9 (penalties)-288.8 (and)-288.9 (weight)-288.8 (matrix)-288.9 (choice.)]TJ /T1_1 1 Tf 22.693 0 Td [(Nucl)-288.8 (Acids)-288.8 (Res)]TJ /T1_0 1 Tf 6.9218 0 Td [(22:4673\226)-119.8 (4680.)]TJ -31.5046 -1.3712 Td [(68.)-499.6 (Guindon)-267.1 (S,)-267.1 (Gascuel)-267.1 (O)-267.1 (\(2003\))-267.1 (A)-267.1 (simple,)-267.1 (fast,)-267.1 (and)-267.1 (accurate)-267.1 (algorithm)-267.1 (to)-267.1 (estimate)-267.1 (large)]TJ 1.8897 -1.3712 Td [(phylogenies)-287 (by)-287 (maximum)-287 (likelihood.)]TJ /T1_1 1 Tf 17.4297 0 Td [(Syst)-286.9 (Biol)]TJ /T1_0 1 Tf 4.1301 0 Td [(52:696)-119.8 (\226704.)]TJ -23.4495 -1.3712 Td [(69.)-499.6 (Strimmer)-291.7 (K,)-291.7 (von)-291.7 (Haeseler)-291.7 (A)-291.7 (\(1996\))-291.7 (Quartet)-291.7 (puzzling:)-291.7 (A)-291.7 (quartet)-291.7 (maximum-likelihood)]TJ 1.8897 -1.3712 Td [(method)-291.7 (for)-291.7 (reconstructing)-291.7 (tree)-291.7 (topologies.)]TJ /T1_1 1 Tf 20.3505 0 Td [(Mol)-291.7 (Biol)-291.7 (Evol)]TJ /T1_0 1 Tf 6.4313 0 Td [(13:964)-119.8 (\226969.)]TJ -28.6715 -1.3712 Td [(70.)-499.6 (Felsenstein)-268.5 (J)-268.5 (\(2002\))-268.5 (PHYLIP)-268.5 (\(Phylogeny)-268.5 (Inference)-268.5 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(y)-352.3 (of)]TJ T* [(ox)-39.8 (ygen)-30 (ic)-259.5 (photosynthetic)-259.5 (machiner)-60 (y)-259.5 (make)19.9 (s)-259.5 (it)-259.5 (dif)-30 (ficult)-259.5 (to)-259.5 (transfer)]TJ T* [(c)-30 (omponents)-448.6 (piecemeal)-448.6 (to)-448.6 (nonphotosynthetic)-448.6 (prok)-30 (ar)-60 (yote)19.9 (s.)-448.6 (In-)]TJ T* [(deed,)-310 (operon)-310 (splitting)-310 (of)-310 (the)-310 (photosynthetic)-310 (apparatus)-310 (require)19.9 (s)]TJ T* [(many)-596.6 (independent)-596.6 (transfers)-596.6 (of)-596.6 (nonc)-30 (ontiguous)-596.6 (operons.)-596.6 (A)-30 (l-)]TJ T* [(though)-219.7 (large-scale)-219.7 (HGT)-219.7 (among)-219.7 (photosynthetic)-219.7 (prok)-30 (ar)-60 (yote)19.9 (s)-219.7 (\(30\))]TJ 0 -1.1124 TD [(may)-189.1 (sugge)19.9 (st)-189.1 (a)-189.1 (c)-30 (omplex)-189.1 (nonlinear)-189.1 (proce)19.9 (ss)-189.1 (of)-189.1 (evolution)-189.1 (that)-189.1 (re)19.9 (sults)]TJ T* [(in)-242.1 (a)-242.1 (mosaic)-242.1 (str)-30 (ucture)-242.1 (of)-242.1 (photosynthetic)-242.1 (pathway)-242.1 (\(60\),)-242.1 (transfer)-242.1 (of)]TJ T* [(the)-296.4 (key)-296.4 (photosynthetic)-296.4 (gene)19.9 (s)-296.4 (are)-296.4 (ver)-60 (y)-296.4 (rare)-296.4 (\(33,)-296.4 (34\).)-296.4 (Transfer)-296.4 (of)]TJ T* [(this)-326.1 (key)-326.1 (pathway)-326.1 (was)-326.1 (only)-326.1 (achieved)-326.1 (by)-326.1 (whole)19.9 (sale)-326.1 (inc)-30 (orporation)]TJ T* [(of)-337.8 (c)-30 (yanobacteria)-337.8 (into)-337.8 (euk)-30 (ar)-60 (yotic)-337.8 (host)-337.8 (cells)-337.8 (\(40)-90 (\226)-90 (44\).)]TJ 1 -1.1124 Td [(Phylogenetic)-628.2 (analysis)-628.2 (of)-628.2 (the)-628.2 (c)-30 (yanobacterial)-628.2 (genome)-628.2 (c)-30 (ore)]TJ -1 -1.1124 Td [(strongly)-624.7 (sugge)19.9 (sts)-624.7 (that)-624.7 (the)-624.7 (last)-624.7 (c)-30 (ommon)-624.7 (ance)19.9 (stor)-624.7 (of)-624.7 (ext)-30 (ant)]TJ T* [(c)-30 (yanobacteria)-325.5 (was)-325.5 (incapable)-325.5 (of)-325.5 (N)]TJ 0 Tc 6.5 0 0 6 183.6219 212.5426 Tm (2)Tj 0.005 Tc 9 0 0 9 189.8322 213.7926 Tm [(fixation)-325.5 (\(Fig.)-325.5 (5\).)-325.5 (This)-325.5 (met)-30 (a-)]TJ -15.3507 -1.1124 Td [(bolic)-421.7 (pathway)-421.7 (appears)-421.8 (to)-421.7 (have)-421.7 (been)-421.7 (acquired)-421.7 (v)-39.8 (ia)-421.7 (HGT)-421.7 (much)]TJ T* [(later)-256.7 (af)-30 (ter)-256.7 (the)-256.7 (origin)-256.7 (of)-256.7 (this)-256.7 (clade,)-256.7 (possibly)-256.7 (as)-256.7 (a)-256.7 (re)19.9 (sult)-256.7 (of)-256.7 (a)-256.7 (\221\221fixed)]TJ T* [(n)-30 (itrogen)-322 (crisis\222\222)-322 (in)-322 (the)-322 (late)-322 (Archean)-322 (and)-322 (early)-322 (Proterozoic)-322 (eons)]TJ T* [(\(61\).)-250.7 (In)-250.7 (this)-250.7 (scenario,)-250.7 (the)-250.7 (ac)-30 (cumulation)-250.7 (of)-250.7 (a)-250.7 (small)-250.7 (c)-30 (oncentration)]TJ T* [(of)-314.1 (ox)-39.8 (ygen)-314.1 (\(re)19.9 (sulting)-314.1 (f)-30 (rom)-314.1 (ox)-39.8 (ygen)-30 (ic)-314.1 (photosynthe)19.9 (sis\))-314.1 (would)-314.1 (have)]TJ T* [(led)-414.2 (to)-414.2 (massive)-414.2 (den)-30 (itrification)-414.2 (of)-414.2 (the)-414.2 (upper)-414.2 (ocean)-414.2 (w)-39.8 (ith)-414.2 (a)-414.2 (c)-30 (on-)]TJ T* [(c)-30 (omit)-30 (ant)-340 (loss)-340 (of)-340 (fixed)-340 (inorgan)-30 (ic)-340 (n)-30 (itrogen)-340 (for)-340 (g)-19.9 (row)-30 (th)-340 (of)-340 (marine)]TJ T* [(photoautotrophs.)-397.8 (This)-397.8 (proce)19.9 (ss)-397.8 (created)-397.8 (an)-397.8 (evolutionar)-60 (y)-397.8 (bottle-)]TJ T* [(neck,)-266.8 (which)-266.8 (potentially)-266.8 (selected)-266.8 (for)-266.8 (the)-266.8 (st)-30 (able)-266.8 (transfer)-266.8 (of)-266.8 (the)]TJ /T1_1 1 Tf 26.3961 0 Td (nif)Tj /T1_0 1 Tf 2.7706 67.9104 Td [(operon)-384.2 (f)-30 (rom)-384.2 (a)-384.2 (\(pre)19.9 (sumably\))-384.2 (heterotrophic)-384.2 (prok)-30 (ar)-60 (yote)-384.2 (to)-384.2 (c)-30 (ya-)]TJ 0 -1.1086 TD [(nobacteria.)-506 (It)-506 (should)-506 (be)-506 (noted)-506 (that)-506 (selection)-506 (of)-506 (the)-506 (specific)]TJ T* [(n)-30 (itrogenase)-618.7 (was)-618.7 (probably)-618.7 (not)-618.7 (related)-618.7 (to)-618.7 (met)-30 (al)-618.7 (availabilit)-30 (y,)]TJ T* [(because)-400.6 (Fe)-400.6 (was)-400.6 (abundant)-400.6 (under)-400.6 (the)19.9 (se)-400.6 (mildly)-400.6 (ox)-49.9 (idizing)-400.6 (c)-30 (ondi-)]TJ T* [(tions)-222.3 (\(62\).)-222.3 (It)-222.3 (was)-222.3 (only)-222.3 (af)-30 (ter)-222.3 (the)-222.3 (\221\221g)-19.9 (reat)-222.3 (ox)-49.9 (idation)-222.3 (event,\222\222)]TJ /T1_2 1 Tf 0 Tc 23.8737 0 Td (\003)Tj /T1_0 1 Tf 0.005 Tc 0.838 0 Td [(2.3)-222.3 (Ga)]TJ -24.7117 -1.1086 Td [(\(1\2263\))-354.5 (and)-354.5 (later,)-354.5 (that)-354.5 (Fe)-354.5 (would)-354.5 (bec)-30 (ome)-354.5 (limiting,)-354.5 (leading)-354.5 (to)-354.5 (the)]TJ T* [(sequential)-524.3 (selection)-524.3 (for)-524.3 (V-)-524.3 (and)-524.3 (Mo-c)-30 (ont)-30 (ain)-30 (ing)-524.3 (n)-30 (itrogenase)19.9 (s.)]TJ T* [(Thus,)-238.4 (under)-238.4 (the)-238.4 (mildly)-238.4 (ox)-49.9 (idizing)-238.4 (c)-30 (onditions)-238.4 (that)-238.4 (prevailed)-238.3 (in)-238.3 (the)]TJ T* [(late)-278.1 (Archean)-278.1 (to)-278.1 (early)-278.1 (Proterozoic,)-278.1 (Fe-based)-278.1 (n)-30 (itrogenase)19.9 (s)-278.1 (would)]TJ T* [(have)-540.4 (been)-540.4 (naturally)-540.4 (selected)-540.4 (w)-39.8 (ithin)-540.4 (the)-540.4 (archaea)-540.4 (and)-540.4 (subse-)]TJ T* [(quently)-225.4 (transferred)-225.4 (to)-225.4 (a)-225.4 (large)-225.4 (g)-19.9 (roup)-225.4 (of)-225.4 (bacteria)-225.4 (v)-39.8 (ia)-225.4 (HGT)-225.4 (\(63\).)-225.4 (In)]TJ T* [(the)-320.7 (late)-320.7 (Proterozoic)-320.7 (and)-320.7 (throughout)-320.7 (the)-320.7 (Phanerozoic,)-320.7 (ox)-39.8 (ygen)-30 (ic)]TJ T* [(photosynthe)19.9 (sis)-373.8 (ultimately)-373.8 (led)-373.8 (to)-373.8 (precipit)-30 (ation)-373.8 (of)-373.8 (insoluble)-373.8 (ox)-49.9 (i-)]TJ T* [(dized)-403.4 (\(ferric\))-403.4 (Fe,)-403.5 (thereby)-403.4 (mak)-30 (ing)-403.4 (this)-403.4 (element)-403.4 (a)-403.4 (major)-403.5 (factor)]TJ T* [(limiting)-418.3 (N)]TJ 0 Tc 6.5 0 0 6 354.762 593.9517 Tm (2)Tj 0.005 Tc 9 0 0 9 361.8075 595.2017 Tm [(fixation)-418.3 (in)-418.3 (the)-418.3 (ocean,)-418.3 (a)-418.3 (c)-30 (ondition)-418.3 (that)-418.3 (appears)-418.3 (to)]TJ -5.2924 -1.1086 Td [(c)-30 (ontinue)-418 (to)-418 (limit)-418 (the)-418 (productiv)-39.8 (it)-30 (y)-418 (in)-418 (the)-418 (c)-30 (ontemporar)-60 (y)-418 (ocean)]TJ T* [(\(64\).)-370 (Macrogenomic)-370 (feature)19.9 (s)-370 (of)-370 (c)-30 (yanobacteria)-370 (potentially)-370 (pro-)]TJ T* [(v)-39.8 (ide)-413.5 (clue)19.9 (s)-413.5 (regarding)-413.5 (the)-413.5 (abilit)-30 (y)-413.5 (of)-413.5 (the)19.9 (se)-413.5 (organ)-30 (isms)-413.5 (to)-413.5 (acquire)]TJ T* [(n)-30 (itrogenase)-445.1 (and)-445.1 (other)-445.1 (gene)19.9 (s.)-445.1 (For)-445.1 (example,)-445.1 (all)-445.1 (c)-30 (yanobacterial)]TJ T* [(diazotrophs)-267.1 (have)-267.1 (sign)-30 (ificantly)-267.1 (larger)-267.1 (genome)19.9 (s)-267.1 (than)-267.1 (their)-267.1 (nonfix-)]TJ T* [(ing)-291.7 (c)-30 (ounterparts)-291.7 (\()]TJ 0 0 1 rg [-0.3 (SI)-291.7 (Table)-291.7 (1)]TJ 0 0 0 rg [-0.2 (\),)-291.7 (sugge)19.9 (sting)-291.7 (that)-291.7 (the)-291.7 (genome)19.9 (s)-291.7 (may)]TJ T* [(have)-223.5 (been)-223.5 (ex)-30 (posed)-223.5 (to)-223.5 (f)-30 (requent)-223.5 (HGT)-223.5 (and)-223.5 (are)-223.5 (more)-223.5 (c)-30 (ompetent)-223.5 (to)]TJ T* [(inc)-30 (orporate)-337.8 (gene)19.9 (s.)]TJ /T1_3 1 Tf -0.01961 Tc 0 -1.9975 TD [(Materials)-313.1 (and)-313.1 (Methods)]TJ 0 Tc 7 0 0 7 314.1761 486.429 Tm [(Gene)-163.6 (Family)-163.6 (Selection.)]TJ /T1_4 1 Tf 10.1048 0 Td [(We)-163.6 (performed)-163.6 (all-against-all)-163.6 (BLAST)-163.6 (\(65\))-163.6 (comparisons)-163.6 (of)]TJ -10.1048 -1.2825 Td [(protein)-305.6 (sequences)-305.6 (for)-305.6 (all)-305.6 (possible)-305.6 (pairs)-305.6 (of)-305.6 (the)-305.6 (13)-305.6 (genomes)-305.6 (of)-305.6 (cyanobacteria)]TJ 0 -1.2825 TD (\()Tj 0 0 1 rg [(SI)-332 (Table)-332 (1)]TJ 0 0 0 rg [(\),)-332 (using)-332 (an)]TJ /T1_5 1 Tf 10.162 0 Td (e)Tj /T1_4 1 Tf 0.888 0 Td [(value)-332 (of)-332 (10)]TJ /T1_2 1 Tf 5 0 0 5 428.4657 470.7244 Tm (\005)Tj /T1_4 1 Tf [-0.1 (4)]TJ 7 0 0 7 437.7354 468.4744 Tm [(as)-332 (a)-332 (lower)-332 (limit)-332 (cutoff,)-332 (and)-332 (reciprocal)]TJ -17.6513 -1.2825 Td [(genome-speci\036c)-430.6 (best)-430.6 (hits)-430.6 (were)-430.6 (identi\036ed.)-430.6 (A)-430.6 (total)-430.6 (number)-430.6 (of)-430.6 (682)-430.6 (protein)]TJ T* [(families)-300.7 (consisting)-300.7 (of)-300.7 (one)-300.7 (gene)-300.7 (per)-300.7 (genome)-300.7 (were)-300.7 (retrieved)-300.7 (and)-300.7 (assigned)-300.7 (to)]TJ T* [(functional)-325.5 (categories)-325.5 (according)-325.5 (to)-325.5 (those)-325.5 (de\036ned)-325.5 (for)-325.5 (the)-325.5 (cluster)-325.5 (of)-325.5 (ortholo-)]TJ T* [(gous)-278 (groups)-278 (\(66\).)]TJ /T1_3 1 Tf 0.00121 Tc 0 -2.9679 TD [(Alignments)-568.6 (and)-568.6 (Tree)-568.6 (Construction.)]TJ /T1_4 1 Tf 16.9821 0 Td [(Protein)-568.6 (sequences)-568.6 (were)-568.6 (aligned)-568.6 (with)]TJ 0 Tc -16.9821 -1.2825 Td [(ClustalW)-373.8 (\(67\),)-373.8 (followed)-373.8 (by)-373.8 (selecting)-373.8 (unambiguous)-373.8 (parts)-373.8 (of)-373.8 (the)-373.8 (alignments)]TJ 0 -1.2825 TD [(excluding)-275.6 (all)-275.6 (gap)-275.6 (sites.)-275.6 (ML)-275.6 (trees)-275.6 (were)-275.6 (computed)-275.6 (with)-275.6 (PHYML)-275.6 (\(68\),)-275.6 (using)-275.6 (the)]TJ T* [(JTT)-159.5 (model)-159.5 (of)-159.5 (substitution)-159.5 (and)-159.5 (the)-159.5 (Gamma)-159.5 (\()]TJ /T1_2 1 Tf [-0.3 (\004)]TJ /T1_4 1 Tf [(\)-based)-159.5 (method)-159.5 (for)-159.5 (correcting)-159.5 (the)]TJ T* [(rate)-200.9 (heterogeneity)-200.9 (among)-200.9 (sites.)-200.9 (Neighbor)-200.9 (joining)-200.9 (\(NJ\))-200.9 (trees)-200.9 (were)-200.9 (constructed)]TJ T* [(by)-295.9 (using)-295.9 (the)-295.9 (distance)-295.9 (matrix)-295.9 (provided)-295.9 (by)-295.9 (TREE-PUZZLE)-295.9 (\(69\))-295.9 (under)-295.9 (a)]TJ /T1_2 1 Tf 31.705 0 Td (\004)Tj /T1_4 1 Tf (-based)Tj -31.705 -1.2825 Td [(model)-342.5 (of)-342.5 (substitution)-342.5 (\(alpha)-342.5 (parameter)-342.5 (estimated,)-342.5 (eight)]TJ /T1_2 1 Tf 27.0712 0 Td (\004)Tj /T1_4 1 Tf 0.9535 0 Td [(rate)-342.5 (categories\))]TJ -28.0247 -1.2825 Td [(and)-314.1 (bootstrapped)-314.1 (by)-314.1 (using)-314.1 (SEQBOOT)-314.1 (and)-314.1 (CONSENSE)-314.1 (from)-314.1 (PHYLIP)-314.1 (\(70\).)-314.1 (See)]TJ 0 0 1 rg /T1_5 1 Tf T* [(SI)-285.7 (Methods)]TJ 0 0 0 rg /T1_4 1 Tf 5.4605 0 Td [(for)-285.7 (concatenation,)-285.7 (consensus,)-285.7 (and)-285.7 (supertree)-285.7 (reconstruction.)]TJ /T1_3 1 Tf -5.4605 -2.9679 Td [(Comparisons)-396.1 (Among)-396.1 (Trees.)]TJ /T1_4 1 Tf 12.7655 0 Td [(Trees)-396.1 (were)-396.1 (compared)-396.1 (with)-396.1 (Treedist)-396.1 (program)-396.1 (in)]TJ -12.7655 -1.2825 Td [(PHYLIP)-406.7 (using)-406.7 (the)-406.7 (branch)-406.7 (score)-406.7 (distance)-406.7 (of)-406.7 (Kuhner)-406.7 (and)-406.7 (Felsenstein)-406.7 (\(71\))-406.7 (to)]TJ T* [(generate)-291.8 (an)]TJ /T1_5 1 Tf 5.9747 0 Td (n)Tj /T1_2 1 Tf 0.9028 0 Td (\006)Tj /T1_5 1 Tf 1.1248 0 Td (n)Tj /T1_4 1 Tf 0.9028 0 Td [(distance)-291.8 (matrix)-291.8 (where)]TJ /T1_5 1 Tf 10.6556 0 Td (n)Tj /T1_4 1 Tf 0.9028 0 Td [(is)-291.8 (the)-291.8 (number)-291.8 (of)-291.8 (trees.)-291.8 (Principal)]TJ -20.4635 -1.2825 Td [(coordinates)-309.3 (analysis)-309.3 (\(PCoA\))-309.3 (was)-309.3 (then)-309.3 (performed)-309.3 (with)-309.3 (the)-309.3 (multidimensional)]TJ T* [(scaling)-275.2 (procedure)-275.2 (in)-275.2 (SAS)-275.2 (software,)-275.2 (Version)-275.2 (8.2)-275.2 (\(SAS)-275.2 (Institute\).)-275.2 (PCoA)-275.2 (allowed)]TJ 0 -1.2841 TD [(us)-251.6 (to)-251.6 (embed)-251.6 (the)]TJ /T1_5 1 Tf 7.7856 0 Td (n)Tj /T1_4 1 Tf 0.8626 0 Td [(trees)-251.6 (in)-251.6 (a)-251.6 (space)-251.6 (of)-251.6 (up)-251.6 (to)]TJ /T1_5 1 Tf 11.2635 0 Td (n)Tj /T1_2 1 Tf 0.8626 0 Td (\005)Tj /T1_4 1 Tf 1.0846 0 Td [(1)-251.6 (dimensions.)-251.6 (By)-251.6 (plotting)-251.6 (the)]TJ -21.859 -1.2841 Td [(objects)-255.3 (\(the)-255.3 (trees\))-255.3 (along)-255.3 (the)-255.3 (most)-255.3 (signi\036cant)-255.3 (two)-255.3 (\036rst)-255.3 (dimensions,)-255.3 (the)-255.3 (major)]TJ T* [(trends)-278 (and)-278 (groupings)-278 (in)-278 (the)-278 (data)-278 (can)-278 (be)-278 (visualized)-278 (graphically.)]TJ 1.2857 -1.2841 Td [(For)-236.6 (each)-236.6 (of)-236.6 (the)-236.6 (682)-236.6 (alignments,)-236.6 (a)-236.6 (comparison)-236.6 (of)-236.6 (the)-236.6 (likelihood)-236.6 (of)-236.6 (the)-236.6 (best)]TJ -1.2857 -1.2841 Td [(topology)-378.3 (with)-378.3 (that)-378.3 (of)-378.3 (the)-378.3 (candidate)-378.3 (topologies)-378.3 (\()]TJ 0 0 1 rg [(SI)-378.3 (Figs.)-378.3 (6)-378.3 (and)-378.3 (7)]TJ 0 0 0 rg [0.1 (\))-378.3 (was)-378.3 (per-)]TJ T* [(formed)-275.2 (with)-275.2 (the)-275.2 (SH)-275.2 (test)-275.2 (\(55\))-275.2 (implemented)-275.2 (in)-275.2 (TREE-PUZZLE.)-275.2 (Similarly,)-275.2 (a)-275.2 (com-)]TJ T* [(parison)-452.5 (of)-452.5 (the)-452.5 (\036ve)-452.5 (backbone)-452.5 (topologies)-452.5 (\(Fig.)-452.5 (2\))-452.5 (was)-452.5 (conducted)-452.5 (with)-452.5 (SH,)]TJ T* [(Kishino\226Hasegawa)-268.7 (\(72\),)-268.7 (and)-268.7 (expected)-268.7 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[(vertically)-207.1 (inherited)-207.1 (indiv)-39.8 (idual)-207.1 (protein-c)-30 (oding)-207.1 (gene)19.9 (s.)-207.1 (But)-207.1 (how)-207.1 (can)]TJ T* [(this)-391.8 (super)-49.9 (ficially)-391.8 (random)-391.7 (gene)-391.8 (transfer)-391.8 (event)-391.8 (ex)-30 (plain)-391.8 (the)-391.8 (c)-30 (on-)]TJ T* [(ser)-60 (ved)-543 (nature)-543 (of)-543 (many)-543 (of)-543 (the)-543 (key)-543 (gene)19.9 (s)-543 (that)-543 (c)-30 (omprise)-543 (the)]TJ T* [(functional)-337.8 (c)-30 (ore)-337.8 (across)-337.8 (all)-337.8 (c)-30 (yanobacterial)-337.8 (t)-30 (axa?)]TJ -0.01489 Tc 1 -1.1111 Td [(A)-30 (l)-0.1 (though)-210.6 (phylogenomic)-210.6 (approache)19.8 (s)-210.6 (are)-210.6 (capable)-210.6 (of)-210.6 (capturing)-210.6 (the)]TJ -1 -1.1111 Td [(c)-30 (onsensus)-174.6 (or)-174.6 (f)-30 (requent)-174.6 (partitions)-174.6 (that)-174.6 (silhouette)-174.6 (the)-174.6 (trend)-174.6 (in)-174.6 (genome)]TJ T* [(evolution,)-482.1 (they)-482.1 (may)-482.1 (not)-482.1 (nece)19.8 (ssarily)-482.1 (guarantee)-482.1 (the)-482.1 (paucit)-30 (y)-482.1 (of)-482.1 (a)]TJ T* [(c)-30 (onf)-90 (licting)-197.3 (phylogenetic)-197.3 (signal)-197.3 (in)-197.3 (genome)-197.3 (space.)-197.3 (The)-197.3 (plural)-197.3 (support)]TJ T* [(for)-200.2 (the)-200.2 (c)-30 (onsensus/supertree/c)-30 (oncatenation)-200.2 (topologie)19.8 (s)-200.2 (indicate)19.8 (s)-200.2 (that)]TJ T* [(the)-427.8 (five)-427.8 (top)-427.8 (topologie)19.8 (s)-427.8 (are)-427.8 (not)-427.8 (sign)-30 (ificantly)-427.8 (dif)-30 (ferent)-427.8 (f)-30 (rom)-427.7 (each)]TJ T* [(other;)-238.7 (that)-238.7 (is,)]TJ /T1_3 1 Tf 0 Tc 5.6908 0 Td (\001)Tj /T1_0 1 Tf -0.01489 Tc 0.8181 0 Td [(90%)-238.7 (of)-238.7 (the)-238.7 (dat)-30 (asets)-238.7 (do)-238.7 (not)-238.7 (discriminate)-238.7 (among)-238.7 (the)]TJ -6.509 -1.1111 Td [(topologie)19.8 (s)-188.5 (\(Fig.)-188.5 (2\).)-188.5 (Do)-188.5 (the)-188.5 (c)-30 (onsensus/supertree/c)-30 (oncatenation)-188.5 (tree)19.8 (s)]TJ T* [(ac)-30 (curately)-308.1 (ref)-90 (lect)-308.1 (organ)-30 (ismal)-308.1 (histor)-60 (y,)-308.1 (or,)-308.1 (on)-308.1 (the)-308.1 (c)-30 (ontrar)-60 (y,)-308.1 (do)-308.1 (they)]TJ T* [(blur)-591.3 (the)-591.3 (vertical)-591.3 (inherit)-30 (ance)-591.3 (signal)-591.3 (by)-591.3 (inc)-30 (orporating)-591.3 (potential)]TJ T* [(HGTs?)-314.1 (There)-314.1 (i)-0.1 (s)-314.1 (a)-314.1 (l)-0.1 (arge)-314.1 (margin)-314.1 (of)-314.1 (uncert)-30 (aint)-30 (y.)-314.1 (Part)-314.1 (of)-314.1 (the)-314.1 (uncer-)]TJ T* [(t)-30 (aint)-30 (y)-406.9 (may)-406.9 (be)-406.9 (due)-406.9 (to)-406.9 (the)-406.9 (strength)-406.9 (of)-406.9 (t)-0.1 (he)-406.9 (Shimodaira\226Hasegawa)]TJ T* [(\(SH\))-335.3 (te)19.8 (st)-335.3 (\(55\),)-335.3 (e)19.8 (s)-0.1 (pecially)-335.3 (when)-335.3 (examin)-30 (i)-0.1 (ng)-335.3 (the)-335.3 (ac)-30 (curac)-30 (y)-335.3 (of)-335.3 (similar)]TJ T* [(topologie)19.8 (s)-0.1 (.)-234.9 (Indeed,)-234.9 (the)-234.9 (SH)-234.9 (te)19.8 (st)-234.9 (was)-234.9 (based)-234.9 (on)-234.9 (the)-234.9 (evaluation)-234.9 (of)-234.9 (only)]TJ T* [(15)-217.2 (of)-217.2 (a)-217.2 (tot)-30 (al)-217.2 (of)-217.2 (13,749,310,575)-217.2 (possible)-217.2 (unrooted)-217.2 (tree)-217.2 (topologie)19.8 (s)-217.2 (for)]TJ T* [(13)-257.3 (specie)19.8 (s)-257.3 (\()]TJ 0 0 1 rg [-0.2 (SI)-257.3 (Figs.)-257.3 (6)-257.3 (and)-257.3 (7)]TJ 0 0 0 rg [-0.3 (\).)-257.3 (A)-30 (lthough)-257.3 (the)-257.3 (majorit)-30 (y)-257.3 (of)-257.3 (the)-257.3 (possible)]TJ T* [(topologie)19.8 (s)-275.3 (would)-275.3 (not)-275.3 (be)-275.3 (supported)-275.3 (by)-275.3 (any)-275.3 (dat)-30 (aset,)-275.3 (t)-0.1 (he)-275.3 (selection)-275.3 (of)]TJ T* [(a)-232.7 (limited)-232.7 (number)-232.7 (of)-232.7 (tree)19.8 (s)-232.7 (may)-232.7 (have)-232.7 (biased)-232.7 (the)-232.7 (analyse)19.8 (s.)-232.7 (But)-232.7 (part)-232.7 (of)]TJ T* [(the)-395.2 (uncert)-30 (aint)-30 (y)-395.3 (can)-395.2 (also)-395.2 (be)-395.2 (attributed)-395.2 (to)-395.2 (the)-395.2 (dat)-30 (a,)-395.3 (most)-395.3 (not)-30 (ably)]TJ T* [(gene)19.8 (s)-318.5 (that)-318.5 (are)-318.5 (subjected)-318.5 (to)-318.5 (HGT)-318.5 (and)-318.5 (homologous)-318.5 (rec)-30 (ombination)]TJ T* [(bet)-30 (ween)-322 (closely)-322 (related)-322 (specie)19.8 (s.)-322 (This)-322 (i)-0.1 (s)-322 (even)-322 (more)-322 (pronounced)-322 (in)]TJ T* [(the)-256 (PCoA,)-256.1 (which)-256 (demonstrate)19.8 (s)-256 (clearly)-256 (that)]TJ /T1_3 1 Tf 0 Tc 17.9433 0 Td (\003)Tj /T1_0 1 Tf -0.01489 Tc 0.8181 0 Td [(53%)-256 (of)-256 (the)-256 (orthologs)]TJ -18.7615 -1.1111 Td [(are)-348.5 (subject)-348.5 (to)-348.5 (HGT)-348.5 (c)-30 (omplicating/diluting)-348.5 (the)-348.5 (vertical)-348.5 (inherit)-30 (ance)]TJ T* [(signal)-317.9 (w)-39.8 (i)-0.1 (thin)-317.9 (the)-317.9 (c)-30 (yanobacterial)-317.9 (phylum)-317.9 (\(Fig.)-317.9 (3\).)]TJ 0.005 Tc 1 -1.1111 Td [(Our)-484.3 (re)19.9 (sults)-484.3 (reveal)-484.3 (that)-484.3 (both)-484.2 (photosynthetic)-484.3 (and)-484.3 (ribosomal)]TJ -1 -1.1111 Td [(gene)19.9 (s)-498.1 (share)-498.1 (similar)-498.1 (evolutionar)-60 (y)-498.1 (historie)19.9 (s)-498.1 (and)-498.1 (belong)-498.1 (to)-498.1 (the)]TJ T* [(c)-30 (yanobacterial)-483 (genome)-483 (c)-30 (ore)-483 (\(Fig.)-483 (3\).)-483 (This)-483 (finding)-483 (of)-483 (limited)]TJ T* [(HGT)-320.7 (in)-320.7 (proteins)-320.7 (w)-39.8 (ith)-320.7 (extraordinarily)-320.7 (c)-30 (onser)-60 (ved)-320.7 (primar)-60 (y)-320.7 (str)-30 (uc-)]TJ T* [(ture)-316.6 (is)-316.6 (c)-30 (onsistent)-316.6 (w)-39.8 (ith)-316.6 (the)-316.6 (c)-30 (omplex)-49.9 (it)-30 (y)-316.6 (hypothe)19.9 (sis;)-316.6 (that)-316.6 (is,)-316.6 (gene)19.9 (s)]TJ T* [(c)-30 (oding)-333.7 (for)-333.7 (large)-333.7 (c)-30 (omplex)-333.7 (systems)-333.7 (that)-333.7 (have)-333.7 (more)-333.7 (macromolec-)]TJ T* [(ular)-306.2 (interactions)-306.2 (are)-306.2 (le)19.9 (ss)-306.2 (subject)-306.2 (to)-306.2 (HGT)-306.2 (than)-306.2 (gene)19.9 (s)-306.2 (c)-30 (oding)-306.2 (for)]TJ T* [(small)-460.3 (assemblie)19.9 (s)-460.3 (of)-460.3 (a)-460.3 (few)-460.2 (gene)-460.3 (products)-460.3 (\(37\).)-460.3 (Translation)-460.3 (in)]TJ T* [(prok)-30 (ar)-60 (yote)19.9 (s)-306.2 (require)19.9 (s)-306.2 (c)-30 (oordinated)-306.2 (assembly)-306.2 (of)-306.2 (at)-306.2 (least)-306.2 (100)-306.2 (gene)]TJ T* [(products,)-403.1 (including)-403.1 (ribosomal)-403.1 (small)-403.1 (and)-403.1 (large)-403.1 (subun)-30 (its,)-403.1 (which)]TJ T* [(interact)-494.7 (w)-39.8 (ith)-494.7 (5S,)-494.7 (16S,)-494.7 (and)-494.7 (23S)-494.7 (rRNA;)-494.7 (numerous)-494.7 (tRNA)-494.6 (and)]TJ T* [(mRNA;)-362.4 (in)-30 (itiation)-362.4 (and)-362.4 (ter)-30 (mination)-362.4 (factors;)-362.4 (ions;)-362.4 (etc.)-362.4 (Similarly,)]TJ T* [(the)-262.7 (ox)-39.8 (ygen)-30 (ic)-262.7 (photosynthetic)-262.7 (apparatus)-262.7 (needs)-262.7 (an)-262.7 (inve)19.9 (stment)-262.7 (of)-262.7 (a)]TJ ET q 157.1738 0 0 283.0328 96.9261 457.8835 cm 1.1111 TL /Im0 Do Q BT 0 0 0 1 k /T1_4 1 Tf 0 Tc 7 0 0 7 51.6761 443.8835 Tm [(Fig.)-243.9 (4.)]TJ /T1_5 1 Tf 3.9889 0 Td [(Frequency)-243.9 (distribution)-243.9 (of)-243.9 (genes)-243.9 (belonging)-243.9 (to)-243.9 (designated)-243.9 (categories)]TJ -3.9889 -1.2857 Td [(within)-157.9 (each)-157.9 (1.0)-157.9 (interval)-157.9 (of)-157.9 (protein)-157.9 (variability.)-157.9 (Protein)-157.9 (variability)-157.9 (was)-157.9 (measured)]TJ 0 -1.2857 TD [(according)-189.9 (to)-189.9 (Rujan)-189.9 (and)-189.9 (Martin\222s)-189.9 (method)-189.9 (\(48\).)-189.9 (Dashed)-189.9 (lines)-189.9 (denote)-189.9 (the)-189.9 (thresh-)]TJ T* [(old)-341.7 (that)-341.7 (segregates)-341.7 (the)-341.7 (predominance)-341.7 (of)-341.7 (distribution)-341.7 (of)-341.7 (genes)-341.7 (in)-341.7 (different)]TJ T* (categories.)Tj ET q 246 0 0 115 52.1562 132.8835 cm 1.2857 TL /Im1 Do Q BT /T1_4 1 Tf 7 0 0 7 51.6761 118.8835 Tm [(Fig.)-286.5 (5.)]TJ /T1_5 1 Tf 4.0741 0 Td [(Phylogenetic)-286.5 (tree)-286.5 (reconstructed)-286.5 (based)-286.5 (on)-286.5 (the)-286.5 (concatenation)-286.5 (of)-286.5 (the)]TJ -4.0741 -1.2857 Td [(323)-184.7 (core)-184.7 (proteins.)-184.7 (The)-184.7 (topology)-184.7 (shown)-184.7 (agrees)-184.7 (with)-184.7 (the)-184.7 (consensus)-184.7 (topology)-184.7 (of)]TJ T* [(the)-429.4 (682)-429.4 (orthologs)-429.4 (\(T3)-429.4 (in)-429.4 (Fig.)-429.4 (2\))-429.4 (and)-429.4 (is)-429.4 (supported)-429.4 (by)-429.4 (almost)-429.4 (all)-429.4 (individual)]TJ T* [(datasets)-320.6 (\(Fig.)-320.6 (2)-320.6 (and)]TJ 0 0 1 rg 9.4516 0 Td [(SI)-320.6 (Fig.)-320.6 (7)]TJ 0 0 0 rg [-0.1 (\).)-320.6 (Bootstrap)-320.6 (probabilities)-320.6 (estimated)-320.6 (by)-320.6 (NJ-)]TJ /T1_3 1 Tf [-0.4 (\004)]TJ /T1_5 1 Tf (/QP/)Tj -9.4516 -1.2857 Td [(ProtML)-250 (with)-250 (1,000)-250 (replications)-250 (are)-250 (shown)-250 (for)-250 (each)-250 (internal)-250 (branch.)-250 (The)-250 (scale)]TJ T* [(bar)-224.4 (refers)-224.4 (to)-224.4 (the)-224.4 (number)-224.4 (of)-224.4 (amino)-224.4 (acid)-224.4 (substitutions)-224.4 (per)-224.4 (site.)-224.4 (The)-224.4 (dashed)-224.4 (line)]TJ T* [(designates)-278 (the)-278 (split)-278 (between)-278 (diazotrophic)-278 (and)-278 (nondiazotrophic)-278 (taxa.)]TJ 6.5 0 0 6.5 51.6761 39.8835 Tm [(Shi)-278 (and)-278 (Falkowski)]TJ 50.639 0 Td (PNAS)Tj /T1_6 1 Tf 9.75 0 0 6.5 403.0802 39.8835 Tm (\001)Tj /T1_4 1 Tf 6.5 0 0 6.5 411.2448 39.8835 Tm [(February)-305.1 (19,)-305.1 (2008)]TJ /T1_6 1 Tf 9.75 0 0 6.5 471.7956 39.8835 Tm (\001)Tj /T1_5 1 Tf 6.5 0 0 6.5 479.9601 39.8835 Tm [(vol.)-305.1 (105)]TJ /T1_6 1 Tf 9.75 0 0 6.5 509.6214 39.8835 Tm (\001)Tj /T1_5 1 Tf 6.5 0 0 6.5 517.786 39.8835 Tm [(no.)-305.1 (7)]TJ /T1_6 1 Tf 9.75 0 0 6.5 539.1335 39.8835 Tm (\001)Tj /T1_4 1 Tf 6.5 0 0 6.5 547.298 39.8835 Tm (2513)Tj /T1_2 1 Tf 0 6.5 -6.5 0 585.1761 81.8949 Tm (EVOLUTION)Tj ET q /GS1 gs 19 0 0 815.5 9 9 cm 1.2857 TL /Im2 Do Q BT 0 g /GS1 gs /T1_7 1 Tf 0 4 -4 0 23 14 Tm (Downloaded by guest on July 10, 2020 )Tj ET endstream endobj 26 0 obj <>stream Adobed         #"""#''''''''''     !! !!''''''''''"  s!1AQa"q2B#R3b$r%C4Scs5D'6Tdt& EFVU(eufv7GWgw8HXhx)9IYiy*:JZjzm!1AQa"q2#BRbr3$4CS%cs5DT 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(s)-334 (in)-334 (the)-334 (shell)-334 (are)-334 (much)-334 (more)-334 (likely)-334 (to)]TJ T* [(be)-337.8 (acquired)-337.8 (v)-39.8 (ia)-337.8 (HGT.)]TJ 1 -1.1326 Td [(We)-229.5 (further)-229.5 (rec)-30 (onstr)-30 (ucted)-229.5 (the)-229.5 (phylogeny)-229.5 (of)-229.5 (the)-229.5 (13)-229.5 (genome)19.9 (s)-229.5 (on)]TJ -1 -1.1326 Td [(the)-537.3 (basis)-537.3 (of)-537.3 (a)-537.3 (superalignment)-537.3 (of)-537.3 (100,776)-537.3 (site)19.9 (s)-537.3 (obt)-30 (ained)-537.3 (v)-39.8 (ia)]TJ T* [(c)-30 (oncatenating)-232.7 (the)-232.7 (323)-232.7 (c)-30 (ore)-232.7 (proteins.)-232.7 (We)-232.7 (used)-232.7 (three)-232.7 (methods,)-232.7 (all)]TJ T* [(of)-241.5 (which)-241.5 (re)19.9 (sult)-241.5 (in)-241.5 (a)-241.5 (tree)-241.5 (hav)-39.8 (ing)-241.5 (the)-241.5 (same)-241.5 (topolog)-39.8 (y)-241.5 (as)-241.5 (that)-241.5 (for)-241.5 (the)]TJ ET q 510.155 0 0 222.6829 51.6761 517.8835 cm 1.1326 TL /Im0 Do Q BT 0 0 0 1 k /T1_4 1 Tf 0 Tc 7 0 0 7 51.6761 503.8835 Tm [(Fig.)-227.3 (3.)]TJ /T1_5 1 Tf 3.9556 0 Td [(PCoA)-227.3 (of)-227.3 (trees)-227.3 (compared)-227.3 (with)-227.3 (topological)-227.3 (distance.)-227.3 (\()]TJ /T1_6 1 Tf [-0.2 (A)]TJ /T1_5 1 Tf [(\))-227.3 (Plot)-227.3 (of)-227.3 (the)-227.3 (two)-227.3 (\036rst)-227.3 (axes)-227.3 (of)-227.3 (the)-227.3 (PCoA)-227.3 (made)-227.3 (from)-227.3 (628)-227.3 (ML)-227.3 (trees.)-227.3 (The)-227.3 (other)-227.3 (54)-227.3 (genes)-227.3 (are)-227.3 (excluded)]TJ -3.9556 -1.2857 Td [(as)-272.3 (a)-272.3 (result)-272.3 (of)-272.3 (axis)-272.3 (demarcation.)-272.3 (The)-272.3 (same)-272.3 (experiment)-272.3 (with)-272.3 (NJ)-272.3 (trees)-272.3 (gave)-272.3 (very)-272.3 (similar)-272.3 (results.)-272.3 (The)-272.3 (ellipse)-272.3 (depicts)-272.3 (323)-272.3 (orthologs)-272.3 (in)-272.3 (the)-272.3 (densest)-272.3 (region)-272.3 (\(t)-1.6 (he)-272.3 (core\))]TJ 0 -1.2857 TD [(of)-200.1 (the)-200.1 (cloud)-200.1 (that)-200.1 (share)-200.1 (a)-200.1 (common)-200.1 (phylogenetic)-200.1 (signal,)-200.1 (whereas)-200.1 (trees)-200.1 (present)-200.1 (in)-200.1 (the)-200.1 (marginal)-200.1 (area)-200.1 (\(the)-200.1 (shell\))-200.1 (are)-200.1 (much)-200.1 (more)-200.1 (likely)-200.1 (to)-200.1 (be)-200.1 (perturbed)-200.1 (by)-200.1 (hori)-0.6 (zontal)]TJ T* [(transfers.)-236.6 (Photosynthetic)-236.6 (genes)-236.6 (are)-236.6 (color)-236.6 (coded)-236.6 (based)-236.6 (on)-236.6 (their)-236.6 (respective)-236.6 (pathways.)-236.6 (Also)-236.6 (shown)-236.6 (are)-236.6 (examples)-236.6 (of)-236.6 (conserved)-236.6 (clusters)-236.6 (of)-236.6 (ribosomal)-236.6 (\(red)-237.9 (text\))-236.6 (and)]TJ T* [(photosynthetic)-243.1 (\(green)-243.1 (text\))-243.1 (genes)-243.1 (that)-243.1 (are)-243.1 (present)-243.1 (in)-243.1 (the)-243.1 (core.)-243.1 (\()]TJ /T1_6 1 Tf [-0.5 (B)]TJ /T1_5 1 Tf [(\))-243.1 (The)-243.1 (PCoA)-243.1 (plotted)-243.1 (against)-243.1 (the)-243.1 (protein)-243.1 (variability.)-243.1 (Protein)-243.1 (variability)-243.1 (was)-243.1 (measured)-243.1 (by)-243.1 (taking)]TJ T* [(the)-221.6 (total)-221.6 (length)-221.6 (of)-221.6 (a)-221.6 (corresponding)-221.6 (tree)-221.6 (as)-221.6 (measured)-221.6 (by)-221.6 (total)-221.6 (amino)-221.6 (acid)-221.6 (substitutions)-221.6 (per)-221.6 (site,)-221.6 (divided)-221.6 (by)-221.6 (the)-221.6 (number)-221.6 (of)-221.6 (sequences)-221.6 (in)-221.6 (the)-221.6 (tree)-221.6 (\(48\).)-221.6 (The)-222.5 (legend)]TJ T* [(is)-278 (the)-278 (same)-278 (except)-278 (that)-278 (photosynthetic)-278 (genes)-278 (are)-278 (collectively)-278 (designated)-278 (as)-278 (green)-278 (dots.)]TJ /T1_4 1 Tf 6.5 0 0 6.5 51.6761 39.8835 Tm (2512)Tj /T1_7 1 Tf 9.75 0 0 6.5 72.1327 39.8835 Tm (\001)Tj /T1_5 1 Tf 6.5 0 0 6.5 80.2973 39.8835 Tm (www.pnas.org)Tj /T1_7 1 Tf [-0.3 (\002)]TJ /T1_5 1 Tf (cgi)Tj /T1_7 1 Tf (\002)Tj /T1_5 1 Tf (doi)Tj /T1_7 1 Tf [-0.1 (\002)]TJ /T1_5 1 Tf (10.1073)Tj /T1_7 1 Tf [-0.1 (\002)]TJ /T1_5 1 Tf (pnas.0711165105)Tj 65.7959 0 Td [(Shi)-278 (and)-278 (Falkowski)]TJ ET q /GS1 gs 19 0 0 815.5 9 9 cm 1.2857 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(exhibit)-280 (their)-280 (own)-280 (un)-30 (ique)]TJ T* [(topologie)19.9 (s.)-306.2 (As)-306.2 (a)-306.2 (re)19.9 (sult,)-306.2 (the)-306.2 (max)-49.9 (imum)-306.2 (number)-306.2 (of)-306.2 (orthologs)-306.2 (that)]TJ T* [(share)-380.4 (a)-380.4 (particular)-380.4 (topolog)-39.8 (y)-380.4 (ac)-30 (c)-30 (ounts)-380.4 (for)-380.4 (only)-380.4 (1.9)-90 (\2262.1%)-380.4 (of)-380.4 (the)]TJ T* [(orthologous)-337.8 (dat)-30 (asets)-337.8 (\(Fig.)-337.8 (1\).)]TJ ET q 196.0473 0 0 152.0147 77.4261 588.8835 cm 1.1149 TL /Im0 Do Q BT 0 0 0 1 k /T1_3 1 Tf 0 Tc 7 0 0 7 51.6761 574.8835 Tm [(Fig.)-224.8 (1.)]TJ /T1_4 1 Tf 3.9507 0 Td [(Distribution)-224.8 (of)-224.8 (tree)-224.8 (topologies)-224.8 (among)-224.8 (682)-224.8 (sets)-224.8 (of)-224.8 (orthologs.)-224.8 (Both)-224.8 (NJ)]TJ -3.9507 -1.2857 Td [(\(black)-340.9 (bars\))-340.9 (and)-340.9 (ML)-340.9 (\(red)-340.9 (bars\))-340.9 (tree)-340.9 (topologies)-340.9 (give)-340.9 (similar)-340.9 (distribution)-340.9 (pat-)]TJ 0 -1.2857 TD [(terns.)-173.3 (There)-173.3 (is)-173.3 (no)-173.3 (unanimous)-173.3 (support)-173.3 (for)-173.3 (a)-173.3 (single)-173.3 (topology;)-173.3 (rather,)-173.3 (most)-173.3 (of)-173.3 (the)]TJ T* [(orthologs)-266.2 (\(58%)-266.3 (and)-266.2 (67%)-266.3 (for)-266.2 (NJ)-266.2 (and)-266.2 (ML)-266.2 (trees,)-266.2 (respectively\))-266.2 (appear)-266.2 (as)-266.2 (single-)]TJ T* [(tons)-278 (that)-278 (associate)-278 (with)-278 (unique)-278 (topologies.)]TJ ET q 386.0957 0 0 258.9168 113.6761 150.8835 cm 1.2857 TL /Im1 Do Q BT /T1_3 1 Tf 7 0 0 7 51.6761 136.8835 Tm [(Fig.)-385.2 (2.)]TJ /T1_4 1 Tf 4.2713 0 Td [(Representative)-385.2 (backbone)-385.2 (tree)-385.2 (topologies.)-385.2 (Phylogenetic)-385.2 (trees)-385.2 (were)-385.2 (constructed)-385.2 (by)-385.2 (using)-385.2 (both)-385.2 (16S)-385.2 (rRNA)-385.2 (gene)-385.2 (and)-385.2 (orthologous)-385.1 (proteins)-385.2 (through)]TJ -4.2713 -1.2857 Td [(phylogenomic)-276.8 (approaches)-276.8 (\(see)]TJ /T1_5 1 Tf 14.6096 0 Td [(Materials)-276.8 (and)-276.8 (Methods)]TJ /T1_4 1 Tf 11.0545 0 Td [(for)-276.8 (details\).)-276.8 (Phylogenetic)-276.8 (tree)-276.8 (construction)-276.8 (methods)-276.8 (are)-276.8 (highlighted)-276.8 (with)-276.8 (colored)-276.8 (horizontal)-276.8 (bars)-276.8 (and)]TJ -25.6641 -1.2857 Td [(text.)-228.9 (Conserved)-228.9 (monophyletic)-228.9 (subgroups)-228.9 (are)-228.9 (shaded.)-228.9 (Row)-228.9 (one)-228.9 (shows)-228.9 (the)-228.9 (proportion)-228.9 (of)-228.9 (orthologs)-228.9 (giving)-228.9 (a)-228.9 (particular)-228.9 (tree)-228.9 (topology)-228.9 (\(NJ,)-228.9 (black)-228.9 (bar;)-228.9 (ML,)-228.9 (re)-1.1 (d)-228.9 (bar\).)]TJ T* [(Also)-284.5 (shown)-284.5 (are)-284.5 (examples)-284.5 (of)-284.5 (proteins)-284.5 (corresponding)-284.5 (to)-284.5 (that)-284.5 (topology.)-284.5 (Row)-284.5 (\036ve)-284.5 (indicates)-284.5 (number)-284.5 (of)-284.5 (datasets)-284.5 (accepting)-284.5 (\()]TJ /T1_5 1 Tf 55.9543 0 Td (Left)Tj /T1_4 1 Tf [(\))-284.5 (or)-284.5 (rejecting)-284.5 (\()]TJ /T1_5 1 Tf [-0.1 (Right)]TJ /T1_4 1 Tf [(\))-284.5 (a)-284.5 (particular)]TJ -55.9544 -1.2857 Td [(topology)-158.7 (in)-158.7 (a)-158.7 (Shimodaira\226Hasegawa)-158.7 (\(SH\))-158.7 (\(55\))-158.7 (test)-158.7 (\()]TJ 0 0 1 rg [-0.3 (SI)-158.7 (Fig.)-158.7 (7)]TJ 0 0 0 rg [(\).)-158.7 (Row)-158.7 (six)-158.7 (shows)-158.7 (the)-158.7 (evaluation)-158.7 (of)-158.7 (the)-158.7 (\036ve)-158.7 (backbone)-158.7 (topologies,)-158.7 (using)-158.7 (the)-158.7 (concatenated)-158.7 (323-core-gene)]TJ T* [(set)-286.9 (\(Fig.)-286.9 (3\))-286.9 (through)-286.9 (Kishino\226Hasegawa)-286.9 (\(72\))-286.9 (\()]TJ /T1_5 1 Tf [-0.5 (Left)]TJ /T1_4 1 Tf [(\),)-286.9 (SH)-286.9 (\(55\))-286.9 (\()]TJ /T1_5 1 Tf [-0.2 (Center)]TJ /T1_4 1 Tf [-0.1 (\),)-286.9 (and)-286.9 (expected)-286.9 (likelihood)-286.9 (weight)-286.9 (\(73\))-286.9 (\()]TJ /T1_5 1 Tf [-0.5 (Right)]TJ /T1_4 1 Tf [(\))-286.9 (tests,)-286.9 (which)-286.9 (infer)-286.9 (a)-286.9 (con\036dence)-286.9 (tree)-286.9 (set.)]TJ /T1_5 1 Tf 70.4425 0 Td (ANA)Tj /T1_4 1 Tf (,)Tj /T1_5 1 Tf -70.4426 -1.2857 Td (Anabaena)Tj /T1_4 1 Tf 5.2547 0 Td [(sp.)-475.7 (PCC7120;)]TJ /T1_5 1 Tf 6.5094 0 Td (AVA)Tj /T1_4 1 Tf (,)Tj /T1_5 1 Tf 2.8647 0 Td [(Anabaena)-475.7 (variabilis)]TJ /T1_4 1 Tf 9.8435 0 Td (ATCC29413;)Tj /T1_5 1 Tf 6.0337 0 Td (CWA)Tj /T1_4 1 Tf (,)Tj /T1_5 1 Tf 3.0867 0 Td [(Crocosphaera)-475.7 (watsonii)]TJ /T1_4 1 Tf 11.2305 0 Td (WH8501;)Tj /T1_5 1 Tf 4.6997 0 Td (GVI)Tj /T1_4 1 Tf (,)Tj /T1_5 1 Tf 2.4207 0 Td [(G.)-475.7 (violaceus)]TJ /T1_4 1 Tf 6.1744 0 Td (PCC7421;)Tj /T1_5 1 Tf 4.7557 0 Td (NPU)Tj /T1_4 1 Tf (,)Tj /T1_5 1 Tf 2.7536 0 Td [(N.)-475.7 (punctiforme)]TJ /T1_4 1 Tf -65.6274 -1.2857 Td (ATCC29133;)Tj /T1_5 1 Tf 5.9059 0 Td (PMM)Tj /T1_4 1 Tf (,)Tj /T1_5 1 Tf 3.0698 0 Td [(P.)-347.8 (marinus)]TJ /T1_4 1 Tf 5.2527 0 Td (MED4;)Tj /T1_5 1 Tf 3.4039 0 Td (PMT)Tj /T1_4 1 Tf (,)Tj /T1_5 1 Tf 2.6818 0 Td [(P.)-347.8 (marinus)]TJ /T1_4 1 Tf 5.2527 0 Td (MIT9313;)Tj /T1_5 1 Tf 4.6279 0 Td (PMS)Tj /T1_4 1 Tf (,)Tj /T1_5 1 Tf 2.6258 0 Td [(P.)-347.8 (marinus)]TJ /T1_4 1 Tf 5.2527 0 Td (SS120;)Tj /T1_5 1 Tf 3.2939 0 Td (SCO)Tj /T1_4 1 Tf (,)Tj /T1_5 1 Tf 2.5148 0 Td [(S.)-347.8 (elongatus)]TJ /T1_4 1 Tf 6.0857 0 Td (PCC7942;)Tj /T1_5 1 Tf 4.6279 0 Td (SYW)Tj /T1_4 1 Tf (,)Tj /T1_5 1 Tf 2.7928 0 Td (Synechococcus)Tj /T1_4 1 Tf 7.1239 0 Td [(sp.)-347.8 (WH8102;)]TJ /T1_5 1 Tf 6.1977 0 Td (SYN)Tj /T1_4 1 Tf (,)Tj /T1_5 1 Tf -70.7101 -1.2857 Td (Synechocystis)Tj /T1_4 1 Tf 6.5111 0 Td [(sp.)-289 (PCC6803;)]TJ /T1_5 1 Tf 6.136 0 Td (TEL)Tj /T1_4 1 Tf (,)Tj /T1_5 1 Tf 2.179 0 Td [(T.)-289 (elongatus)]TJ /T1_4 1 Tf 6.024 0 Td (BP-1;)Tj /T1_5 1 Tf 2.623 0 Td (TER)Tj /T1_4 1 Tf (,)Tj /T1_5 1 Tf 2.29 0 Td [(Trichodesmium)-289 (erythraeum)]TJ /T1_4 1 Tf 13.1361 0 Td (IMS101.)Tj 6.5 0 0 6.5 51.6761 39.8835 Tm [(Shi)-278 (and)-278 (Falkowski)]TJ 50.639 0 Td (PNAS)Tj /T1_6 1 Tf 9.75 0 0 6.5 403.0802 39.8835 Tm (\001)Tj /T1_3 1 Tf 6.5 0 0 6.5 411.2448 39.8835 Tm [(February)-305.1 (19,)-305.1 (2008)]TJ /T1_6 1 Tf 9.75 0 0 6.5 471.7956 39.8835 Tm (\001)Tj /T1_4 1 Tf 6.5 0 0 6.5 479.9601 39.8835 Tm [(vol.)-305.1 (105)]TJ /T1_6 1 Tf 9.75 0 0 6.5 509.6214 39.8835 Tm (\001)Tj /T1_4 1 Tf 6.5 0 0 6.5 517.786 39.8835 Tm [(no.)-305.1 (7)]TJ /T1_6 1 Tf 9.75 0 0 6.5 539.1335 39.8835 Tm (\001)Tj /T1_3 1 Tf 6.5 0 0 6.5 547.298 39.8835 Tm (2511)Tj /T1_0 1 Tf 0 6.5 -6.5 0 585.1761 81.8949 Tm (EVOLUTION)Tj ET q /GS1 gs 19 0 0 815.5 9 9 cm 1.2857 TL /Im2 Do Q BT 0 g /GS1 gs /T1_7 1 Tf 0 4 -4 0 23 14 Tm (Downloaded by guest on July 10, 2020 )Tj ET endstream endobj 3 0 obj <>stream H [kA4Q. &jdU &=g1_qz.0 0ϳTutMϼnT46=3T^/`xwFGb~d`LH^f6e55hufo-tMtFȾzbYҗY(b5+A4gvU5rk9{F*FZFK-(vĸ#G'!NBr8r"lzO;-+^s2qkx%g5f-IoƷlJ&Wұ%CoFP?c[JVK G9U*Cۣ[*G0w%S#rw5t.C$ QWS50Q,P|?#6^C҅0UG;u)Mj8 S)L|@3aKCtJfj!MB!<&.4/*Gd8&t VYCi9.WىSK¼tD9981*ʥfW z0Ys*[DƆr$mNtcQDȹ+"gksLhDd?IN41hٚa@@xRZ.s,wgS+$oG=qnM0 px z.DTό.n)̞a@Bڵt(NQ>19.=j4'\B-ki>D_Η?gafHW[iUӎ63a .0 .0 .0 .0 .0 x䘂8rh^wa?5=o^oe8~J<}l@0T=٧Fd4 tO Q &U€wa iL0cY`T/b &&? 0`RW?gw 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